Professor William Sherwin
Emeritus Professor
Field of Research: 
Molecular ecology, biodiversity, population genetic models
Contact details:
+61 2 9385 2119

Room 522, D26 Building
UNSW, Kensington 2052


Reseach Students


Oliver Manlik (PhD candidate - dolphin MHC, fitness, and population Viability)

Sandra Vogel (PhD candidate  - penguin MHC and population Viability)

Claire Brandenburger (PhD candidate  - weed invasion and speciation)

Ryan Shofner (PhD candidate  - Macadmia Lacebug geneflow and species identity)



Jackie Chan (PhD 2015) Prawn Genetics

Anna Kopps (PhD 2011) Dolphin genetics and social behaviour, computer modelling.

Maria Cardoso (PhD 2011) Quoll conservation Genetics

Celine Frere (PhD 2009) Dolphin genetics and social behaviour.

Lee Ann Rollins (PhD 2008) Starling invasion genetics and mitochondrial selection

Maio, Gianluca  (MSc 2009)  Modelling genetic consequences of asymmetrical dispersal betwen populations

Clare Holleley (PhD 2008) Genetic estimation of dispersal, and effects of selection (in Drosophila)

Angela  Penn (PhD 2006)  Vertebrate Population Viability Analysis

Michael Kruetzen (PhD 2003) Dolphin genetics and social behaviour

Melissa Gunn (PhD 2003) Quantitative and population genetics of small populations (in Drosophila)

Deryn Alpers (PhD 1996)  Wombat genetics and population viability

Andrea Taylor (PhD 1994) Wombat conservation genetics



BIOS2021   Genetics

BIOS6671  Biodiversity and Conservation of Natural Resources




89.  Chao  A, Jost L, Hsieh TC, Ma KH, Sherwin WB, Rollins LA. 2015.  Expected Shannon entropy and Shannon differentiation between subpopulations for neutral genes under the finite island model.  PLoS ONE (accepted 14 Apr 2015 PONE-D-14-34101R1)

88. Rollins LA, Whitehead MR, Woolnough A, Sinclair R, Sherwin WB. 2015. Is there evidence of selection in the dopamine receptor D4 gene in Australian invasive starling populations.  Current Zoology 61:505-519  Accepted 2 April 2015.  CZ-2015-0010.R2

87.  Sherwin WB,  Frommer M, Sved J, Raphael K, Oakeshott JG, Shearman DCA, Gilchrist ASG. 2015.  Tracking invasion and invasiveness in Queensland fruit flies – from classical genetics to ‘omics. Current Zoology  61:477-487.

86. Chao A, Jost L, Hsieh TC, Ma KH, Sherwin WB Rollins LA.  (provisional acceptance 27 Nov 2014)  Expected Shannon entropy and Shannon differentiation between subpopulations for neutral genes under the finite island model. PLoS ONE PONE-D-14-34101

85.  Gilchrist AS, Shearman DCA, Frommer M, Raphael KA, Deshpande NP, Wilkins MR, Sherwin WB, Sved JA. 2014. The draft genome of the pest tephritid fruit fly Bactrocera tryoni: resources for the genomic analysis of hybridising species.  BMC Genomics 15:1153.

84.  Raphael KA, Shearman DCA, Gilchrist S, Sved JA, Morrow JL, Sherwin WB, Riegler M, Frommer M. 2014  Australian endemic pest tephritids: genetic, molecular and microbial tools for improved Sterile Insect Technique.  BMC Genetics 15(Suppl 2):S9  doi:10.1186/1471-2156-15-S2-S9

83. Chan JT, SherwinWB, Taylor MD.  (2014 accepted) "A tool for tracking genetic contributions of wild Penaeus (Melicertus) plebejus broodstock to hatchery populations.  Animal Genetics accepted 29 April 2014 Manuscript ID AnGen-14-01-0028

82.  Cardoso MJ, Mooney N, Eldridge MDB, Firestone KB, Sherwin WB. 2014. Genetic monitoring reveals significant population structure in eastern quolls – implications for the conservation of a threatened carnivorous marsupial.  Australian Mammalogy   36:169-177.

81.  Kopps AM, Ackermann C, Sherwin WB, Allen SJ, Bejder L, Krützen, M.  2014 accepted.  Cultural transmission of tool use combined with habitat specialisations leads to fine-scale genetic structure in bottlenose dolphins.  Proceedings of the Royal Society B.  281:  20133245; DOI: 10.1098/rspb.2013.3245. Published 19 March 2014

80.  Kopps AM, KrützenM, Allen SJ, Bacher K, Sherwin WB.  2013 Characterizing the socially transmitted foraging tactic ‘sponging’ by bottlenose dolphins (Tursiops sp.) in the western gulf of Shark Bay, Western Australia. Marine Mammal Science30(3): 847-863. DOI: 10.1111/mms.12089

79.  Rollins LA, Moles AT, Buswell JM, Lam S, Brandenburger CR, Buitenwerf R, Flores-Moreno H, Nielsen KB, Couchman E, Brown GS, Thomson FJ, Hemmings F, Frankham R & Sherwin WB.  2013.  High genetic diversity is not essential for successful introduction.   Ecology and Evolution3:4501-4517. doi/10.1002/ece3.824/pdf

78.  Chan JT, Waqairatua S, Appleyard S, Preston N, Dixon TJ, Sherwin WB, Taylor MD 2013.  Novel polymorphic microsatellite loci for the eastern king prawn, Penaeus (Melicertus) plebejus. Conservation Genetics Resources 5:1125-1128  COGR-D-13-00196

77  Holleley CE,  Nichols RA, Whitehead MR, Adamack AT, Gunn MR, Sherwin WB. 2014   Testing single-sample estimators of effective population size in genetically structured populations.  Conservation Genetics 15:23-35.  DOI 10.1007/s10592-013-0518-3

76 Cristescu R, Cahill V, Sherwin WB, Handasyde K,  Carlyon K, Whisson D, Herbert CQ, Carlsson BLJ, Wilton AN, Cooper DW. 2012.   Corrigendum to: Inbreeding and testicular abnormalities in a bottlenecked population of koalas (Phascolarctos cinereus).  Wildlife Research 39:374 - 374

75 Kopps AM, SherwinWB. 2012.  Modelling the emergence and stability of a vertically transmitted cultural trait in bottlenose dolphins, Animal Behaviour 84: 1347–1362.     

ERA A JIF 5yr 3.5911yr 3.493

74  Randic S, Connor RC, Sherwin WB, Krützen M. 2012 A novel mammalian social structure in Indo-Pacific bottlenose dolphins (Tursiops sp.): complex male-male alliances in an open social network.  Proceedings of the Royal Society B  279, 3083–3090.

ERA A* JIF 5yr 5.4 1yr 5.0

73  Thurlby KAG, Wilson PG, Sherwin WB, Connelly C, Rossetto M. 2012. Reproductive bet-hedging in a rare yet widespread rainforest tree, Syzygium paniculatum (Myrtaceae).  Austral Ecology 37: 936–944.

ERA B JIF 5 yr 2.0, 1 yr 1.8

72  Cannell BBL, Pollock KHJ, Bradley SJS, Wooller RD, Sherwin WB, Sinclair J. 2011 accepted.  Augmenting mark-recapture with beach counts to estimate the abundance of little penguins on Penguin Island, Western Australia.  Wildlife Research 38:491 – 500.

ERA B JIF 5 yr 1.3, 1 yr 1.2

71  Dewar RC, Sherwin WB, Thomas E, Holleley CE, Nichols RA. 2011. Predictions of single-nucleotide polymorphism differentiation between two populations in terms of mutual information.  Molecular Ecology  20:3156–3166.

ERA A JIF 5 yr 6.6, 1 yr 6.0

70  Rollins LA, Woolnough AP, Sinclair R, Mooney NJ, Sherwin WB.  2011. Mitochondrial DNA offers unique insights into invasion history of the common starling.  Molecular Ecology 20:2307–2317.

ERA A JIF 5 yr 6.6, 1 yr 6.0

69 Holleley CE, Nichols RA, Whitehead MR, Gunn MR, Gupta J, Sherwin WB.  2011. Induced dispersal in wildlife management: experimental evaluation of the risk of hybrid breakdown and the benefit of hybrid vigor in the F1 generation.  Conservation Genetics 12: 31-40.

ERA B JIF 5yr 1.9, 1yr 1.3

68 **  Frère CH,  Mann J, Krützen M, Connor RC, Bejder L, Sherwin WB. 2010

Nature and nurture: a step towards investigating their interactions in the wild. Communicative & Integrative Biology - Addendum Article (Accepted 25 November 2010) Volume 4, Issue 2   March/April 2011

Not ERA listed

67 Frère CH, Krützen M, Mann J, Connor R, Bejder L, Sherwin B. 2010.  Social and genetic interactions drive fitness variation in a free-living dolphin population

Proceedings of the National Academy of the USA 107:19949-19954.

ERA A* JIF 5yr 10.6 1yr 9.7

66   Connor RC, Watson-Capps JJ, Sherwin WB, Krutzen M. 2010. A new level of complexity in the male alliance networks of Indian ocean bottlenose dolphins (Tursiops sp.) Biology Letters 7: 623-626

ERA A JIF 5yr 3.6 1yr 3.8

65  Sherwin WB.  Review: Entropy and Information Approaches to Genetic Diversity and its Expression: Genomic Geography.  Entropy 2010, 12(7): 1765-1798; doi:10.3390/e12071765

(in Special Issue “Entropy in Genetics and Computational Biology”)

ERA C  JIF 1 yr 1.1

64  Frère CH, Krützen M, Mann J, Watson-Capps JJ, Tsai YJ, Patterson EM, Connor R, Bejder L, Sherwin WB. 2010.  Home range overlap, matrilineal and biparental kinship drive female associations in bottlenose dolphins.  Animal Behaviour 80: 481-486.

ERA A JIF 5yr 3.1 1yr 3.4

63 Frere C, Krützen M, Ward P, Kopps AM, Mann J, Sherwin WB.  2010.  Inbreeding tolerance and fitness costs in wild bottlenose dolphins.  Proceedings of the Royal Society B 277:2667-2673.

ERA A* JIF 5yr 5.4 1yr 5.0

62 **  Sherwin WB.  2010. Mechanisms of Evolution.  Chapter 33, pp783-808 in Knox, B, Ladiges P, Evans B, Saint R (eds)  Biology: an Australian Focus 4th Ed, McGraw-Hill North Ryde.

Textbook, Not ERA listed

61 Valsecchi E, Corkeron PJ, Galli P, Sherwin W, Bertorelle G.  2010 Genetic evidence for sex-specific migratory behaviour in the western South Pacific humpback whales.  Marine Ecology Progress Series 398: 275–286

ERA A JIF 5yr 3.0 1yr 2.5

60 Cristescu R, Sherwin WB, Handasyde K, Cahill V, Cooper DW.  2010.  Detecting bottlenecks using BOTTLENECK 1.2.02 in wild populations: the importance of the microsatellite structure.  Conservation Genetics 11:1043-1049.

ERA B JIF 5yr 1.9, 1yr 1.3

59 Cristescu R Cahill V Sherwin WB Handasyde K Carlyon K Whisson D Herbert CA Carlsson BLJ Wilton A Cooper D. 2009 Inbreeding and testicular abnormalities in a bottlenecked population of koalas (Phascolarctos cinereus).  Wildlife Research 36:299-308 ERA B JIF 5yr 1.3, 1yr 1.2

58 Rollins LA, Woolnough AP, Wilton AN, Sinclair R and Sherwin WB.  2009.  Invasive species can’t cover their tracks: using microsatellites to assist management of starling (Sturnus vulgaris) populations in Western Australia.  Molecular Ecology 18: 1560–1573 ERA A JIF 5 yr 6.6, 1 yr 6.0

57 Cardoso MJ  Eldridge MDB  Oakwood M Rankmore B  Sherwin WB Firestone KB.  2009. Effects of founder events on the genetic variation of translocated island populations: implications for conservation management of the northern quoll. Conservation Genetics 10:1719-1733. ERA B JIF 5yr 1.9, 1yr 1.3

56  Rossetto M, Kooyman R, Sherwin WB, Jones R. 2008.  Dispersal limitations, rather than bottlenecks or habitat specificity, can restrict the distribution of rare and endemic rainforest trees.  American Journal of Botany  95(3): 321–329.

ERA A JIF 5yr 3.5 1yr 3.0

55 ** Holleley CE Sherwin WB.  2007.  Two robust multiplex reactions for high-throughput microsatellite genotyping in Drosophila melanogaster.  Drosophila Information Service 90:140-144.

Not ERA listed

54 Sherwin WB and Graves JAM. 2006.  What Marsupials can do for Genetics and what Genetics can do for Marsupials. In Armati PJ, Dickman, CR. and Hume ID (Eds.) Marsupials.  Cambridge Univ. Press, Cambridge.

Book, Not ERA listed

53 Barclay SD, Costello B. Sherwin WB. 2006. Limited cross-species microsatellite amplification and the isolation and characterization of new microsatellite markers for the greater stick-nest rat (Leporillus conditor). Molecular Ecology Notes 6:882-885

Renamed Molecular Ecology Resources, ERA C

52 Rollins L-A, Woolnough A, Sherwin WB.  2006   Population genetic tools for pest management: A review.  Wildlife Research 33: 251 – 261
ERA B JIF 5 yr 1.3 1 yr 1.2

51 Sherwin WB, Jabot F, Rush R,  Rossetto M 2006 Measurement of biological information with applications from genes to landscapes  Molecular Ecology15:2857-2869.

ERA A JIF 5 yr 6.6, 1 yr 6.0

50 KrützenM, MannJ, HeithausM, ConnorRC, BejderL & SherwinWB. 2005. Cultural Transmission of Tool Use in Bottlenose Dolphins.  Proceedings of the National Academy of the USA 102: 8939-8943

ERA A* JIF 5yr 10.6 1yr 9.7

49  Valsecchi E, Amos W, Raga JA, Podestà M, William Sherwin WB.   2004.  The effects of inbreeding on mortality during a morbillivirus outbreak in the Mediterranean striped dolphin (Stenella coeruleoalba)    Animal Conservation 7: 139-146

ERA B  JIF 5yr 3.0 1 yr 2.9

48  Krützen M, Connor RC, Mann J, Barré LM and Sherwin WB. (2004).  “O father where art thou? – Paternity assessment in an open fission-fusion society of wild bottlenose dolphins (Tursiops sp) in Shark bay, WA.  Molecular Ecology 13:1975-1990

ERA A JIF 5 yr 6.6, 1 yr 6.0

47  Krützen MK, Sherwin WB, Berggren P, and Gales NJ. (2004). Population structure in an inshore cetacean revealed by microsatellite and mtDNA analysis: bottlenose dolphins (Tursiops sp) in Shark Bay, Western Australia. Marine Mammal Science 20:28-47


46 ** Sherwin WB.  2003.  Conserving Genetic Information.  IN: Quested, S (Editor)  50 Years of DNA 1953 - 2003  Business Weekly / The Wellcome Trust (London / Cambridge).

Book Not ERA listed

45 Alpers DL, Taylor AC, Banks SC, Sunnucks P, Bellman SA and Sherwin WB.  2003 Pooling hair samples to increase DNA yield for PCR.  Conservation Genetics 4:779-788

ERA B JIF 5yr 1.9, 1yr 1.3

44  Penn AM, Sherwin WB, Lunney D and Banks P.  (2003).  The effects of a low-intensity fire on small mammals and lizards in a logged, burnt forest.  Wildlife Research30:477-486

ERA B JIF 5 yr 1.3 1 yr 1.2

43  Krüetzen M, Sherwin WB, Connor RC, Barré LM, van de Casteele T, Mann J and Brooks R. 2003  Contrasting relatedness patterns in bottlenose dolphins (Tursiops sp.) with different alliance strategies.  Proceedings of the Royal Society B 270:497-502

ERA A* JIF 5yr 5.4 1yr 5.0

42  Kruetzen M, Barre LM, Heithaus MR, Simms C, Moller LM and Sherwin WB.  2002.  A biopsy system especially for small cetaceans: darting success and wound healing in Tursiops sp.  Marine Mammal Science 18: 863-878


41.  Krützen M, Valsecchi EA, Connor RC and Sherwin WB.  2002.  Characterization of microsatellite loci in Tursiops aduncus. Molecular Ecology Notes 1: 170-172.

Renamed Molecular Ecology Resources, ERA C

40 **   Sherwin WB 2002.  A lesson to us all - “The last Tasmanian Tiger: The history and extinction of the Thylacine”, By Robert Paddle  Quarterly Review of Biology 77:179-180

Book Review – NOT peer reviewed


39.  Lunney D, O’Neill L, Matthews A and Sherwin WB 2002.  Modelling mammalian extinction and forecasting recovery: koalas at Iluka (NSW Australia). Biological Conservation 106: 101-113

ERA A  JIF 5yr 4.0 1yr 3.5

38. Krützen M, Valsecchi E, Connor RC, Sherwin WB.  2001.  Characterization of microsatellite loci in Tursiops aduncusMolecular Ecology Notes 1 (3), 170-172

37 Firestone KB, Houlden BA, Sherwin WB, and Geffen E.  2000.  Variability and differentiation of microsatellites in the genus Dasyurus and conservation implications for the large Australian carnivorous marsupials.  Conservation Genetics 1: 115-133.

ERA B JIF 5yr 1.9, 1yr 1.3

36. Sherwin WB and Moritz CM (2000).  Managing and monitoring genetic erosion.  Pp9-34  In Young A and Clarke G (eds).  Genetics, Demography and the viability of fragmented populations.  Cambridge University Press, Cambridge.

Book, Not ERA listed

35. Sherwin WB, Timms P, Wilcken J, and Houlden B.  2000.  Analysis and Conservation Implications of Koala Genetics.  Conservation Biology 14:639-649.

ERA A*  JIF 5yr 6.0 1 yr 4.9

34. Penn AM, Sherwin WB, Gordon G, Lunney D, Melzer A and Lacy RC. 2000  Demographic Forecasting in Koala Conservation.  Conservation Biology 14:629-638.

ERA A*  JIF 5yr 6.0 1 yr 4.9

33. Firestone K Elphinstone MS Sherwin WB and Houlden BA. 1999.  Phylogeographic population structure of tiger quolls Dasyurus maculatus (Dasyuridae: marsupialia), an endangered carnivorous marsupial.  Molecular Ecology 8:1613-1626

ERA A JIF 5 yr 6.6, 1 yr 6.0

32. Fowler EV, Houlden BA, Sherwin WB, Hoeben P, and Timms P.  1998.  Genetic variation in captive koalas (Phascolarctos cinereus): parentage determination and individual identification.  Biochemical Genetics: 36:193-204.

31. Luikart G. Sherwin WB, Steele, BM and Allendorf FW. 1998. Usefulness of Molecular Markers for detecting population bottlenecks via monitoring genetic change.  Molecular Ecology 7(8):963-974.

ERA A JIF 5 yr 6.6, 1 yr 6.0

30. Luikart G, Allendorf FW and Sherwin WB. 1998.  Distortion of allele frequency distributions provides a test for recent population bottlenecks.  Journal of Heredity   89(3):238-247.

ERA B  JIF 5yr 2.3

29. Hoyle SD, Alpers D and Sherwin WB.  1998.  A population simulation of the northern hairy-nosed wombat Lasiorhinus krefftii for conservation and management.pp 165-179 in Wells RT and Pridmore PA (eds) Wombats  Surrey Beatty and Sons Sydney

28. Alpers D, Taylor AC and Sherwin WB.  1998  Genetic structure of populations of the southern hairy-nosed wombat Lasiorhinus latifrons. pp 192-197 in Wells RT and Pridmore PA (eds) Wombats Surrey Beatty and Sons Sydney

27. Taylor AC, Alpers D and Sherwin WB.  1998.  Remote censusing of northern hairy-nosed wombats Lasiorhinus krefftii via genetic typing of hairs collected in the field.  pp 156-164 in Wells RT and Pridmore PA (eds) Wombats Surrey Beatty and Sons Sydney

26. Brown A, Young A, Burdon J, Christidis L, Clarke G, Coates D and Sherwin WB.  1997.  Genetic indicators for state of the environment reporting.  Australia: state of the environment technical paper series (environmental indicators), Department of the Environment, Sport, and territories, Canberra.

25. Luikart G, Painter J, Crozier R, Westerman M and Sherwin WB.  1997.  Characterization of microsatellite loci in the endangered long-footed potoroo Potorous longipes.  Molecular Ecology 6: 497-498.

ERA A JIF 5 yr 6.6, 1 yr 6.0

24. Taylor AC, Horsup A, Johnson CN, Sunnucks P  and Sherwin WB.  1997.  Relatedness structure detected by microsatellite analysis and attempted pedigree construction in an endangered marsupial, the northern hairy-nosed wombat, Lasiorhinus krefftii.  Molecular Ecology 6(1): 9-19. ERA A JIF 5 yr 6.6, 1 yr 6.0

23. Taylor AC, Marshall Graves JA, Murray ND, OBrien SJ, Yuhki N) and Sherwin WB.  1997.  Conservation genetics of the koala (Phascolarctos cinereus):  Low mitochondrial DNA variation amongst southern Australian populations.  Genetical Research Cambridge 69: 25-33. ERA C

22. Moritz C,  Sherwin W, and Limpus CJ  (1996)  Applications of genetics to the conservation management of Australian fauna.  pp. 442-456  In Smith, TB and Wayne, RK et al (eds)  Molecular Genetic Approaches to Conservation Oxford Univ. Press, New York

21 Houlden BA, Greville WD and Sherwin WB.  1996. Evolution of MHC class I loci in marsupials: Characterization of sequences from koala (Phascolarctos cinereus) (vol 13, pg 1124, 1996) Molecular Biology and Evolution 13: 1407-1407.


20. Houlden BA, Greville WD and Sherwin WB.  1996.  Evolution of MHC class I loci in marsupials: characterisation of sequences from koala (Phascolarctos cinereus).  Molecular Biology and Evolution 13:1119-1127.


19. Houlden BA, England PR, Taylor AC, Greville WD and Sherwin WB.  1996.  Low genetic variability of the koala (Phascolarctos cinereus) in south eastern Australia following a severe population bottleneck.  Molecular Ecology 5:269-281.   

ERA A JIF 5 yr 6.6, 1 yr 6.0

18. Houlden BA, England P and Sherwin WB.  1996.  Paternity exclusion in koalas using hypervariable microsatellites.  Journal of Heredity87:149-152.

ERA B  JIF 5yr 2.3

17. Moritz C, Coates DJ, Sherwin WB, Clancy T. Limpus .CJ. 1994. Population ecology and genetics. In Conservation biology in Australia and Oceania: 359–362, Vol. xii. Moritz C. & Kikkawa J. (Eds). Chipping Norton: Surrey Beatty

16. Taylor AC, Sherwin WB, and Wayne RK, 1994. The use of simple sequence loci to measure genetic variation in bottlenecked species: the decline of the northern hairy-nosed wombat (Lasiorhinus krefftii)  Molecular Ecology  3:277-290                   

ERA A JIF 5 yr 6.6, 1 yr 6.0

15. Robinson NA, Sherwin WB, and Murray ND).  1993  Use of VNTR loci to reveal population structure in the eastern barred bandicoot, Perameles gunnii.  Molecular Ecology  2:195-207

ERA A JIF 5 yr 6.6, 1 yr 6.0

14 **  Sherwin, W. B.  (1993 - not refereed scientifically).  Can small remnants have a role in genetically relevant conservation? - workshop report.  pp 83-85  In:  Howard, J.L., and Rawling, J. Proceedings of the 1992 Urban Bushland Conference  Kuringai Council, Sydney.

13. Taylor AC and Sherwin WB.  1992.  Wombat microsatellites in time and space.  Ancient DNA Newsletter 1:33

12. Sherwin WB 1992.  Conservation of Genetic Resources.  pp171-192 in Dixon PI (ed)  Population Genetics and its Applications to Fisheries Management and Aquaculture Centre for Marine Science, University of NSW, Sydney.

11. Taylor AC, Marshall Graves JA, Murray ND and Sherwin WB.  1991.  Conservation genetics of the koala (Phascolarctos cinereus) II. Limited variability in minisatellite DNA sequences. Biochemical Genetics 29: 355-363.


10. Sherwin WB, Murray ND, Graves JAM and Brown PR.  1991.  Measurement of genetic variation in endangered populations: Bandicoots (Marsupialia: Peramelidae) as an example. Conservation Biology 5: 103-108.

ERA A*  JIF 5yr 6.0 1 yr 4.9

9. Sherwin WB.  1991.  Collecting mammalian tissue and data for genetic studies. Mammal Review 21: 21-30. ERA B

8. Robinson NA, Sherwin WB and Brown PR.  1991.  A note on the status of Tasmanian Eastern Barred Bandicoot, Perameles gunnii.  Wildlife Research 18:451-7 ERA B

7. Sherwin WB and Murray ND. 1990.  Population and conservation genetics of marsupials.  Australian Journal of Zoology 37: 161-180. ERA B

6 .Sherwin WB and Brown PR.  1990.  Problems in the estimation of the effective size of a population of the Eastern Barred Bandicoot, Perameles gunnii, at Hamilton, Victoria.  pp 367-374  In: Seebeck, J.H., Brown, P.R., Wallis, R.H., and Kemper, C.M. (eds.) Bandicoots and Bilbies.  Surrey Beatty and Sons, Sydney.

5. Sherwin WB  1990. Post-translational modification of Alpha-1 Antitrypsin and Alkaline Phosphatase in the marsupial Perameles gunnii. Biochemical Genetics 28: 111-115. ERA C

4 ** Sherwin WB.  (1990 – not refereed scientifically).  Genetic implications of reduced fertility.  9pp.  IN: Dept Conservation and Environment, Vic (ed)  Proceedings of the fertility control in wildlife conference 12-24 Nov 1990.  Atkinson Conference Services, East Melbourne.

3. Robinson NA, Sherwin WB, Murray ND and Graves JAM.  1990   Applications of conservation genetics to the Eastern Barred Bandicoot, Perameles gunnii.  pp 109-131  In: Clark T and Seebeck JH (eds.) Management of small populations. Chicago Zoological Society, Brookfield, Ill, USA.

2. Sherwin WB, Murray ND, Graves JAM and Brown PR.  (1989).  Minimum research on conservation genetics of wild populations. In: Hicks, M., and Eiser, P. (eds.) The conservation of threatened species and their habitats.  Occasional Paper No. 2.  Australian Committee of the International Committee for the Conservation of Nature and Natural Resources; Sydney.

1. **  Sherwin WB.  1989.  Conservation genetics and taxonomy of  Perameles gunnii. In: Brown, P.R. Management plan for the conservation of the Eastern Barred Bandicoot, Perameles gunnii, in Victoria. NPWD, ARIER, Victorian Ministry for Conservation, Forests, and Lands, Tech. Rep. Ser. No. 63.